ELATERIDARIUM

Přijato 01.09.2014

ELATERIDARIUM ročník 8 (2014) strany 120-142 (27.10.2014)

První údaje o sedmi druzích kovaříků (Coleoptera, Elateridae) a přehled druhů, známých z území Mongolska

Andrea JARZABEK-MÜLLER & Tamás NÉMETH

1 Anton-Hilz-Strasse 42, D-94566 Riedlhütte, Bavaria, Germany
e-mail: andrea_jarzabek@yahoo.de
2 Hungarian Natural History Museum, H-1088 Budapest, Baross utca 13, Hungary
e-mail: haesito@gmail.com

Abstract. First records of Ampedus gagatinus Candèze, 1895, Cardiophorus discicollis Herbst, 1806, Denticollis flavipes Germar, 1846, Eanus costalis costalis Paykull, 1800, Fleutiauxellus maritimus Curtis, 1840, Negastrius nadezhdae Dolin, 1971, Oedostethus latissimus Cherepanov, 1957 and a complete checklist of Elateridae of Mongolia are presented with notes on distribution, morphological characteristics and photos of habitats for the new species and the endemic Cardiophorus kaszabi Gurjeva, 1969.

Key Words: Coleoptera, Elateridae, Ampedus gagatinus, Cardiophorus discicollis, Cardiophorus kaszabi, Denticollis flavipes, Eanus costalis costalis, Fleutiauxellus maritimus, Negastrius nadezhdae, Oedostethus latissimus, characteristics, distribution, new records, Palearctic, Mongolia, habitat

Introduction

     The fauna of the family Elateridae (Coleoptera) in Mongolia is currently inadequately known. Previous publications on the Mongolian elaterid beetles exist by Fleutiaux (1936), Gurjeva (1968, 1969, 1971, 1972, 1974, 1975a, b, 1976, 1980 and 1984), Emetz et al. (1974), Tegahzhargal (1989) and Bussler (2013). Currently 94 species of Elateridae have been recorded from Mongolia. The country is dominated by desert (Gobi) and steppe habitats, which are less suitable for the elaterids more associated with forest and riparian habitats. Due to the traditionally low intensity of forest use in Mongolia, these southern parts of the boreal forest belt in the Palaearctic provide natural habitats with many species, which are rare in Europe, but occurring here in strong populations (Müller et al. 2013). The appearance of international wood companies are new threat to these extensive virgin forests, important habitat for elaterids.
     Elaterids were collected during three trips in 2010, 2012 and 2014 covering the region of Ulaanbaatar (surrounding), Changai, Gobi, Altai and the Forests of the Khan Khentii area between Zuuncharaa and Khonin Nuga (buffer zone of the Strictly Protected Area of Khan Khentii (Mühlenberg et al. 2011)). In addition, the click-beetles of the collecting trips of the eminent Hungarian coleopterist Dr. Zoltán Kaszab (1915–1986) between 1963 and 1968 in Mongolia and deposited in the collection of the Hungarian Natural History Museum, Budapest, Hungary (HNHM) were examined.
     In this contribution seven species are recorded for the first time in Mongolia.

First records and some comments on the characteristics

Subfamily Elaterinae Leach, 1815
Tribe Ampedini Gistel, 1856

Ampedus gagatinus Candèze, 1895
Elater compactus Candèze, 1891
Elater candezei Kraatz, 1879
(Fig. 1-4)

Material examined. 1 ♀ – Mongolia, Selenge Prov., West-Khentii, Khan Khentii SPA, 5 km W of Khonin Nuga Research Station, 1370 m NN a. s. l. (N 49.11787° E 107.26859°), 30.VII.2010, A. Jarzabek- Müller leg. et coll., A. Jarzabek-Müller det. 2010; 1 ♂ – Mongolia, Selenge Prov., West-Khentii, Khan Khentii SPA, Khonin Nuga Research Station, 919 m NN a. s. l. (N 49.08748° E 107.29216°), 31.VII.2010, J. Müller leg., A. Jarzabek-Müller det. 2010, A. Jarzabek-Müller coll.
Collection circumstances: The both specimen were found under the bark of a large burned dead larch (Larix sibirica) in a Dark Taiga Forest of the upper montane belt with Pinus sibirica and in a Larix sibirica-Betula platyphylla Sub Taiga Forest (Fig. A1, A2).
Characteristics: Ampedus gagatinus Candèze differs from A. aethiops Lacordaire by the larger and more robust body. The pronotum is strongly punctured but shiny. Elytral striae are strongly impressed. The interstriae are convex to strongly convex near elytra base (Fig. 1, 2). Legs and antennae are brown-reddish.
A. gagatinus also differs from the similar species - A. cognatus Gurjeva, A. fulvipes Motschulsky and A. pallipes Kraatz - by the umbilicated punctures along the lateral margin of pronotum (punctures along the lateral margin of pronotum in the back third of these species are simple).
Length: 12 - 16 mm, Sternite (Fig. 3), Aedeagus (Fig. 4).

Distribution: Russia (East Siberia, West Siberia, Far East), Kazakhstan (Cate 2007), China (Liaoning) (Platia & Gudenzi 2006), China (Heilongjiang) (Mertlik & Cooter 2007).

First record for Mongolia. E. L. Gurjeva has already suspected this species in northern Mongolia (Gurjeva 1975b).


Fig. 1. Ampedus gagatinus Candèze, ♂ (from Mongolia) Body length 13.4 mm	Fig. 2. Ampedus gagatinus Candèze, ♀ (from Mongolia) Body length 15.7 mm

Fig. 3. A. gagatinus Cand., ♂ 9th and 10th sternite	Fig. 4. A. gagatinus Cand., ♂ Aedeagus

Subfamily Denticollinae Stein & J. Weise, 1877
Tribe Denticollini Stein & Weise, 1877

Denticollis flavipes Germar, 1846
Campylus flavipes Germar, 1846
(Fig. 5)

Material examined. 1 ♂ – Mongolia, Selenge Prov., West-Khentii, Khan Khentii SPA, Khonin Nuga Research Station, 978 m NN a. s. l. (N 49.08757° E 107.29188°), 17.VI.2012, A. Jarzabek-Müller leg. et coll,, A. Jarzabek-Müller det. 2014
Collection circumstances: This species was taken by sweeping of herbage in Larix sibirica-Betula platyphylla Sub Taiga Forest (Fig. A13).
Length: 9 - 13 mm.
Distribution: Russia (Far East) (Cate 2007).
First records for Mongolia.

Fig. 5. Denticollis flavipes Germar, ♂ and aedeagus (from Mongolia)
Body length 11.5 mm

Tribe Ctenicerini Fleutiaux, 1936

Eanus costalis costalis Paykull, 1800
Diacanthus parvicollis Mannerheim, 1853
(Fig. 6-11)

Material examined. 1 ♂ – Mongolia, Selenge Prov., West-Khentii, Khan Khentii SPA, 8 km W of Khonin Nuga Research Station, 1541 m NN a. s. l. (N 49.16406° E 107.30070°), 19.VI.2012, A. Gruppe leg., A. Jarzabek-Müller det. 2014, A. Jarzabek-Müller coll.; 1 ♀ – Mongolia, Selenge Prov., West-Khentii, Khan Khentii SPA, Khonin Nuga Research Station, 960 m NN a. s. l. (N 49.08731° E 107.31025°), 21.VI.2012, J. Müller leg., A. Jarzabek-Müller det. 2014, A. Jarzabek-Müller coll.
Collection circumstances: This species was taken by sweeping of fir (Abies sibirica) and spruce (Picea obovata) canopy in Larix sibirica-Betula platyphylla Sub Taiga Forest and riverine Forest (Fig. A3).
Characteristics: Eanus costalis costalis Paykull (Subgenus Pareanus) is distinguished from E. guttatus Germar and E. singularis Mannerheim (Subgenus Eanus) by the larger size, the flattened frontal collar margin and the characteristic genital (Fig. 8-11).
Length: 6.5 - 10 mm, Aedeagus (Fig. 8, 9).
Distribution: Finland, Norway, Sweden, Russia (North European Territory, East Siberia, Far East, West Siberia), Nearctic Region (Cate 2007).
First records for Mongolia.


Fig. 6. Eanus costalis costalis Paykull, ♂ (from Mongolia) Body length 7.8 mm	Fig. 7. Eanus costalis costalis Paykull, ♀ (from Mongolia) Body length 8.8 mm


Fig. 8. Aedeagus of E. costalis costalis Payk. (from Mongolia)	Fig. 9. Aedeagus of E. costalis costalis Payk. (Gurjeva, 1989)	Fig. 10. Aedeagus of E. guttatus Germ. (Gurjeva, 1989)	Fig. 11. Aedeagus of E. singularis Mann. (Gurjeva, 1989)

Subfamily Negastriinae Nakane & Kishii, 1956

Fleutiauxellus maritimus Curtis, 1840
Cryptohypnus gracilis Mulsant & Guillebeau, 1855
Cryptohypnus morio Kiesenwetter, 1858
Cryptohypnus scotus Candèze, 1860
(Fig. 12-14)

Material examined. 5 ♂ – Mongolia, Zavakhan Prov., 30 km N Uliastai, 2149 m NN a. s. l. (N 47.90016° E 097.04172°), 09.VI.2012, A. Jarzabek-Müller leg. et coll., A. Jarzabek-Müller det. 2014
Collection circumstances: This species was found under stones along the bank of a partially dried up small mountain river (Fig. A4).
Characteristics: The sides of the last sternite are a little emarginated before apex, which ceased in a tip (Fig. 13). The 2th antennal segment is globose and as dull as the following segments. The scutellum is clearly longer than wide.
The similar species Fleutiauxellus algidus Sahlberg differs from F. maritimus Curtis by the simple rounded 5th sternite. In F. algidus the second antennal segment is tapered or cylindric and shiny. The scutellum is not or slightly longer than wide.
Length: 4 - 6 mm, Sternite (Fig. 13), Aedeagus (Fig. 14).
Distribution: Austria, Czech Republic, Finland, France, Germany, Great Britain, Italy, Norway, Poland, Slovakia, Slovenia, Sweden, Switzerland, Ukraine (Cate 2007)
First record for Mongolia.


	Fig. 13. 5th sternite of F. maritimus Cur., ♂ (from Mongolia)
Fig. 12. Fleutiauxellus maritimus Curtis, ♂ (from Mongolia) Body length 4.5 mm - 4.8 mm
	Fig. 14. Aedeagus of F. maritimus Cur. (from Mongolia)

Oedostethus latissimus Cherepanov, 1957
(Fig. 15)

Material examined. 1 ♀ – Mongolia, Uvs Prov., Somon Öndörchangaj, 1900 m, 11.VII.1968, Z. Kaszab leg., E. L. Gurjeva det., HNHM coll.
Length: 5.3-6 mm (Fig. 15).

Distribution: Russia (East Siberia) (Cate, 2007).
First record for Mongolia.

Fig. 15. Oedostethus latissimus Cherepanov, ♀ (from Mongolia)
Body length 6 mm

Negastrius nadezhdae Dolin, 1971
(Fig. 16-20)

Material examined. 4 ♂, 4 ♀ – Mongolia, Selenge Prov., West-Khentii, Khan Khentii SPA, Khonin Nuga Research Station, River Yeröö Gol, 919 m NN a. s. l. (N 49.08748° E 107.29216°), 28.VII.-08.VIII.2010 and 20.VI.2012; 5 ♂, 13 ♀, 1 Larva – Mongolia, Khovd Prov., Altai, near Bayan-Enger, 1799 m NN a. s. l.
(N 48.42078° E 090.97356°), 06.VI.2012, A. Jarzabek-Müller leg. et coll., A. Jarzabek-Müller det. 2014; 2 ♂ – Mongolia, Bayan-Olgii Prov., Altai, Ölgii, Tsagaannuur, 1669 m NN a. s. l. (N 49.06376° E 090.18032°), 07.VI.2012, A. Jarzabek-Müller leg. et coll., A. Jarzabek-Müller det. 2014.
Collection circumstances: This species was found under stones or running on sand along sandbanks of mountain rivers (Fig. A5-A8).
Characteristics: Negastrius nadezhdae Dolin is similar to N. pulchellus Linnaeus but of this distinguished by the rugosed, matted and punctured elytral intervals (Fig. 19). The marginal punctures of several first elytral intervals of N. pulchellus form continuous rows and interstices are nearly smooth and shiny (Fig. 21).
N. nadezhdae differs from N. arenicola Boheman by the shallower elytral intervals. The elytral striae of N. arenicola are more deeply engraved, especially in the anterior half of elytra (Fig. 22).
Length: 2.7 – 4.2 mm, Aedeagus (Fig. 20).
Distribution: Kirghizia (East Than-Shan Mountains) (Dolin 1971) (Fig. 23), Nearctic (North America: U.S.A. (Massachusetts, New Hampshire)) (Wells 1996).
First records for Mongolia.


Fig. 16. Negastrius nadezhdae Dolin, ♂ (from Mongolia) Body length 2.9 mm - 3.8 mm	Fig. 17. N. nadezhdae Dolin, ♀ (from Mongolia)	Fig. 18. Dolin's drawing of N. nadezhdae, ♂


Fig. 19. Elytral intervals and striae of N. nadezhdae Dol., ♂ (from Mongolia)	Fig. 20. Aedeagus of N. nadezhdae Dol. (from Mongolia)


Fig. 21. Elytral intervals and striae of N. pulchellus L., ♂ (from Bulgaria)	Fig. 22. Elytral intervals and striae of N. arenicola Boh., ♂ (from Poland)

Fig. 23. Known Distribution of Negastrius nadezhdae Dolin in the Palearctic Region

Subfamily Cardiophorinae Candèze, 1860
Tribe Cardiophorini Candèze, 1860

Cardiophorus discicollis Herbst, 1806
Cardiophorus brevinotaticollis Pic, 1913
Cardiophorus ganglbaueri Buysson, 1897
Cardiophorus pleuralis Buysson, 1899
(Fig. 24)

Material examined. 1♀– Mongolia, Chovd Prov., Mongol Altaj Gebirge, Uljasutajn gol, 45 km NNO von Somon Bulgan, 1400 m, 6.VII.1966, Z. Kaszab leg., E. L. Gurjeva det. 1967, HNHM coll.
Length: 6 –7.5 mm (Fig. 24).

Distribution: Austria, Bosnia Herzegovina, Bulgaria, China (Xinjiang), Croatia, Czech Republic, Germany, Georgia, Greece, Hungary, Italy, Israel, Kasakhstan, Macedonia, Moldavia, Poland, Romania, Russia (Central and South European Territory, West Siberia), Serbia and Montenegro, Slovakia, Slovenia, Switzerland, Syria, Tajikistan, Turkey, Ukraine (Cate, 2007).
First record for Mongolia.

Fig. 24. Cardiophorus discicollis Herbst, ♀ (from Mongolia)
Body length 7 mm

Notes on the endemic species Cardiophorus kaszabi Gurjeva, 1969
(Fig. 25–29, 31)

Material examined. 2 ♂ – Mongolia, Khovd Prov., Altai, Khovd, near Myangad, 1263 m NN a. s. l. (N 48.15238° E 091.74115°), 05.VI.2012, A. Jarzabek-Müller leg. et coll., A. Jarzabek-Müller det. 2014
Collection circumstances: This species was collected from shrubs of Caragana pygmaea (Fig. A15).
Characteristics: Cardiophorus kaszabi Gurjeva is similar to C. przewalskii Gurjeva and C. vexillarius Candèze. However, the species can be distinguished as follows:
The punctures of pronotum of C. kaszabi are the same size (Fig. 29), while die punctures of pronotum of C. vexillarius are two different sizes (Fig. 30).
The shape of scutellum of C. kaszabi is heart-shaped and slightly longer than wide (Fig. 31), whereas the scutellum of C. przewalskii is narrow and clearly longer than wide (Fig. 32).
Length: 6.8 – 8 mm, Aedeagus (Fig. 27, 28)
Distribution: Mongolia (Ömnögovi Province (South Gobi), Dundgovi Province (Middle Gobi)) (Gurjeva 1969, Cate 2007).


Fig. 25. Cardiophorus kaszabi Gurjeva, ♂; 1st specimen (from Mongolia) Body length 6.9 mm	Fig. 26. Cardiophorus kaszabi Gurjeva, ♂; 2st specimen (from Mongolia) Body length 7.7mm	Fig. 27. Aedeagus of C. kaszabi Gurj. (from Mongolia)	Fig. 28. Gurjeva's drawing of aedeagus of C. kaszabi (Gurjeva, 1969)

Fig. 29. Punctures of pronotum of C. kaszabi Gurj. (from Mongolia)	Fig. 30. Gurjeva's drawing of the punctures of pronotum of C. vexillarius Cand. (Gurjeva, 1966)	Fig. 31. Scutellum of C. kaszabi Gurj. (from Mongolia)	Fig. 32. Gurjeva's drawing of scutellum of C. przewalskii Gurj. (Gurjeva, 1966)

List of species of Elateridae of Mongolia
with some habitat photos

Family Elateridae Leach, 1815
Subfamily Agrypninae Candèze, 1857
Genus Danosoma Thomson, 1859

Danosoma conspersa Gyllenhal, 1808
Danosoma fasciata Linnaeus, 1758 (Fig. A9, A10)

Genus Lacon Laporte, 1838

Lacon altaicus Candèze, 1882

Genus Aeoloderma Fleutiaux, 1928

Aeoloderma savioi Fleutiaux, 1936

Genus Aeoloides Schwarz, 1906

Aeoloides grisescens Germar, 1844
Aeoloides hauseri Reitter, 1896

Subfamily Elaterinae Leach, 1815
Genus Agriotes Eschscholtz, 1829

Agriotes lineatus Linnaeus, 1767
Agriotes meticulosus Candèze, 1863
Agriotes nadezhdae Cherepanov, 1965
Agriotes obscurus Linnaeus, 1758
Agriotes rugipennis Schwarz, 1891
Agriotes sericatus Schwarz, 1891
Agriotes soederbomi Fleutiaux, 1936
Agriotes sputator Linnaeus, 1758
Agriotes squalidus squalidus Schwarz, 1891
Agriotes unicolor Koenig, 1889
Agriotes zinovjevi Gurjeva, 1967

Genus Dalopius Eschscholtz, 1829

Dalopius marginatus Linnaeus, 1758
Dalopius puerilis Candèze, 1879

Genus Ampedus Dejean, 1833

Ampedus balteatus Linnaeus, 1758
Ampedus basalis Mannerheim, 1852
Ampedus bocakorum Schimmel, 2003
Ampedus gagatinus Candèze, 1895 New record for Mongolia (Fig. A1, A2)
Ampedus nigrinus Herbst, 1784
Ampedus pallipes Kraatz, 1879
Ampedus pomonae Stephens, 1830
Ampedus pomorum Herbst, 1784
Ampedus sanguineus Linnaeus, 1758
Ampedus sanguinolentus sanguinolentus Schrank, 1776
Ampedus sobrinus Motschulsky, 1860

Genus Sericus Eschscholtz, 1829

Sericus brunneus brunneus Linnaeus, 1758

Genus Astanchus Gurjeva, 1979

Astanchus ussuriensis Gurjeva, 1975

Subfamily Melanotinae Candèze, 1859
Genus Melanotus Gurjeva, 1979

Melanotus mongolicus Gurjeva, 1968

Subfamily Hypnoidinae Schwarz, 1906
Genus Berninelsonius Leseigneur, 1970

Berninelsonius hyperboreus Gyllenhal, 1827

Genus Hypnoidus Dillwyn, 1829

Hypnoidus koltzei Reitter, 1910
Hypnoidus rivularius rivularius Gyllenhal, 1808

Genus Ligmargus Stibick, 1976

Ligmargus depressus Gebler, 1847 (Fig. A11)

Subfamily Pleonominae Semenov & Pjatakova, 1936
Genus Pleonomus Ménétriés, 1849

Pleonomus canaliculatus Faldermann, 1835

Subfamily Denticollinae Stein & Weise, 1877
Genus Cidnopus Thomson, 1859

Cidnopus koltzei Reitter, 1895 (Fig. A16)
Cidnopus parallelus Motschulsky, 1860 (Fig. A16)

Genus Pheletes Kiesenwetter, 1858

[Pheletes quercus Olivier, 1790]
Pheletes lythrodes Germar, 1813: According to Reitter (1905, p. 19) it also present in northern Mongolia, but Gurjeva (1972) revised this statement; distribution only in Central and Southern Europe.
Pheletes reitteri Gurjeva, 1976

Genus Denticollis Piller & Mitterpacher, 1783

Denticollis borealis Paykull, 1800 (Fig. A13)
Denticollis cinctus Candèze, 1863 (Fig. A13)
Denticollis flavipes Germar, 1846 New record for Mongolia (Fig. A13)
Denticollis linearis Linnaeus, 1758
Denticollis mongolicus Motschulsky, 1860
Denticollis nigricollis Gebler, 1830
Denticollis varians varians Germar, 1846

Genus Diacanthous Reitter, 1905

Diacanthous undulatus DeGeer, 1774 (Fig. A13)

Genus Megathous Reitter, 1905

Megathous dauricus Mannerheim, 1852
Megathous sedakovii Mannerheim, 1852

Genus Anostirus Thomson, 1859

Anostirus boeberi Germar, 1842 (Fig. A11, A12)
Anostirus castaneus castaneus Linnaeus, 1758

Genus Ctenicera Latreille, 1829

Ctenicera cuprea Fabricius, 1775

Genus Eanus LeConte, 1861

Eanus costalis costalis Paykull, 1800 New record for Mongolia (Fig. A3)

Genus Liotrichus Kiesenwetter, 1858

Liotrichus affinis Fabricius, 1775

Genus Hypoganomorphus Dolin, 1975

Hypoganomorphus laevicollis Mannerheim, 1852 (Fig. A13)

Genus Metanomus Buysson, 1887

Metanomus infuscatus Eschscholtz, 1829

Genus Paraphotistus Kishii, 1966

Paraphotistus impressus impressus Fabricius, 1792

Genus Poemnites Buysson, 1894

Poemnites hamirensis Cherepanov, 1957

Genus Prosternon Latreille, 1834

Prosternon montanum Gurjeva, 1980
Prosternon sericeum Gebler, 1824 (Fig. A14)

Genus Pseudanostirus Dolin, 1964

Pseudanostirus altaicus altaicus Eschscholtz, 1829
Pseudanostirus ecarinatus Stepanov, 1930
Pseudanostirus vicinus Gurjeva, 1984

Genus Selatosomus Stephens, 1830

Selatosomus (Pristilophus) melancholicus melancholicus Fabricius, 1798
Selatosomus (Pristilophus) punctatissimus Ménétriés, 1851
Selatosomus (Selatosomus) aeneus Linnaeus, 1758
Selatosomus (Selatosomus) centralis Candèze, 1882
Selatosomus (Selatosomus) confluens confluens Gebler, 1830
Selatosomus (Selatosomus) coreanus Miwa, 1928 (Fig. A11)
Selatosomus (Selatosomus) latus Fabricius, 1801
Selatosomus (Selatosomus) songoricus Kraatz, 1879 (Fig. A11)

Subfamily Negastriinae Nakane & Kishii, 1956

Genus Fleutiauxellus Méquignon, 1930

Fleutiauxellus maritimus Curtis, 1840 New record for Mongolia (Fig. A4)

Genus Negastrius Thomson, 1859

Negastrius nadezhdae Dolin, 1971 New record for Mongolia (Fig. A5-A8)
Negastrius pulchellus Linnaeus, 1761

Genus Neohypdonus Stibick, 1971

Neohypdonus arcticus altaicus Cherepanov, 1957

Genus Oedostethus LeConte, 1853

Oedostethus aerarius Reitter, 1895
Oedostethus graniger Cherepanov, 1956 (Fig. A8)
Oedostethus kaszabi Gurjeva, 1968
Oedostethus latissimus Cherepanov, 1957 New record for Mongolia!
Oedostethus mediocris Gurjeva, 1972
Oedostethus mystax Gurjeva, 1971
Oedostethus varians Gurjeva, 1968 (Fig. A8)

Genus Zorochros Thomson, 1859

Zorochros hummeli Fleutiaux, 1936
Zorochros murinoides Gurjeva, 1963 According to Gurjeva (1975b) it is also present in southern Mongolian Altai Mts.

Subfamily Cardiophorinae Candèze, 1860

Genus Cardiophorus Eschscholtz, 1829

Cardiophorus discicollis Herbst, 1806 New record for Mongolia
Cardiophorus ebeninus Germar, 1824 (Fig. A16-A17)
Cardiophorus gebleri gebleri Candèze, 1860
Cardiophorus kaszabi Gurjeva, 1969 (Fig. A15)
Cardiophorus keyserlingi Koenig, 1889
Cardiophorus vestigialis Erichson, 1840
Cardiophorus vulgaris Motschulsky, 1860

Genus Dicronychus Brullé, 1832

Dicronychus subulipennis Faldermann, 1835

Genus Paracardiophorus Schwarz, 1895

Paracardiophorus erythrurus Candèze, 1882
Paracardiophorus kaszabi Gurjeva, 1968
Paracardiophorus pullatus pullatus Candèze, 1873 (Fig. A8)

Acknowledgments

We thank Michael Mühlenberg and Enkhmaa Ayush for intensive support during field sampling in Khonin Nuga and Heinz Bussler, Volker Dorka, Dieter Dotschkal, Axel Gruppe, Gisela Merkel-Wallner, Jörg and Paul Müller for support during sampling. In addition, we thank Ottó Merkl for text review.

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Appendix: Habitat photos