Anton-Hilz-Strasse 42, D-94566 Riedlhütte, Bavaria, Germany
e-mail: andrea_jarzabek@yahoo.de

Abstract. The article provides some corrections and additions to the list of Elaterids of Mongolia. Oedostethus sincerus Gurjeva, 1975 and Oedostethus petrenkoi Dolin, 1992 are new for Mongolia. The announcement of Fleutiauxellus maritimus Curtis, 1840 for Mongolia has been revised. An identification key to the species of the genus Oedostethus LeConte in Mongolia is presented.

Zusammenfassung. Der Artikel ergänzt und korrigiert die kürzlich erschienene Liste der Elateriden der Mongolei. Oedostethus sincerus Gurjeva, 1975 und Oedostethus petrenkoi Dolin, 1992 sind neu für die Mongolei. Die Meldung über den Neunachweis von Fleutiauxellus maritimus Curtis, 1840 für die Mongolei wird revidiert. Ein Bestimmungsschlüssel der aus der Mongolei bekannten Oedostethus-Arten wird präsentiert.

Key Words: Coleoptera, Elateridae, Aeolosomus rossii, Agriotes soederbomi, Agriotes unicolor, Ampedus basalis, Ampedus mannerheimi, Ampedus tristis, Fleutiauxellus maritimus, Oedostethus aerarius, Oedostethus graniger, Oedostethus kaszabi, Oedostethus latissimus, Oedostethus mediocris, Oedostethus mystax, Oedostethus petrenkoi, Oedostethus sincerus, Oedostethus varians, Oedostethus, identification key, male, new record, distribution, Palearctic, Mongolia
 

Introduction

     In a recent publication, a list of Mongolian Elaterids has been presented (Jarzabek-Müller & Neméth, 2014). This list is now revised and complemented. The revision shows that the species Fleutiauxellus maritimus Curtis, recorded from the province Zavakhan in Mongolia has been mistaken. The species is Oedostethus latissimus Tsherepanov. The confusion is based on identical traits for both genera Fleutiauxellus Méquignon and Oedostethus LeConte. It seems obvious that the current Oedostethus genus has to be further divided. Some species probably have to be transferred to Fleutiauxellus, and to other, maybe new genera. Japanese authors (Kishii 1976, 1987, Ohira 1988) described many new genera for Japanese Negastriinae, but outside Japan similar species, still are described as Oedostethus, despite clear distinction in their aedeagus. Beyond Oedostethus latissimus Tsherepanov also Oedostethus mystax Gurjeva from Mongolia and Sibiria or Oedostethus submontanus Dolin & Bessolitzina recorded from Sibiria show characteristics typical for Fleutiauxellus. Thus further research based on phylogenetic is required to illuminate the relationship of this complex in elaterids.
     Moreover, the revision revealed a specimen determined by Gurjeva as Oedostethus latissimus (Jarzabek-Müller & Neméth, 2014), as a female of Oedostethus petrenkoi Dolin.
     To facilitate the determination of the species of the genera Oedostethus from Mongolia under consideration of Fleutiauxellus maritimus a key using information from Russian, English and German literature has been developed. (Dolin 1992, Gurjeva 1968, 1969, 1975a, b, Reitter 1895, 1905, Tsherepanov 1957).  
 

Complementary notes on the list of Elateridae of Mongolia

Aeolosomus rossii Germar, 1844
subfamily Agrypninae Candèze, 1857

E. L. Gurjeva reported this species for Mongolia already in 1968 (Gurjeva 1968, 1969).

Agriotes unicolor Koenig, 1889
subfamily Elaterinae Candéze, 1863

On the basis of type material Agriotes soederbomi Fleutiaux, 1936 was synonymized with Agriotes unicolor Koenig, 1889 by G. Platia (2007).

Ampedus mannerheimi Suzuki, 2013
subfamily Elaterinae Candéze, 1863

W. Suzuki renamed Ampedus basalis (Mannerheim, 1852) as Ampedus mannerheimi Suzuki, 2013.

Ampedus tristis Linnaeus, 1758
subfamily Elaterinae Candéze, 1863

1915 G. G. Jacobson mentioned this species for Mongolia (Jacobson 1905-1915). However, this information has been second-guessed.

Fleutiauxellus maritimus Curtis, 1840
(Fig. 5, 16, 34, 53)
subfamily Negastriinae Nakane & Kishii, 1956

This species has to be removed from the checklist of Mongolia.

Oedostethus petrenkoi Dolin, 1992
(Fig. 2, 31, 49)
subfamily Negastriinae Nakane & Kishii, 1956

E. L. Gurjeva determined the specimen in Fig. 31 in 1968 as Oedostethus latissimus Tsherep., but it is a female of Oedostethus petrenkoi Dolin, a species first described in 1992 by Dolin (Dolin 1992). The females of both species are very similar, but the antennae of Oedostethus petrenkoi almost extend to posterior angles of pronotum, while the antennae of Oedostethus latissimus extend two segments past posterior angles of pronotum. The elytra of Oed. petrenkoi are more convex, while the elytra of Oed. latissimus are more flat.

Material examined. 1 ♀ – Mongolia, Uvs Prov., Somon Öndörchangaj, 1900 m, 11.VII.1968, Z. Kaszab leg., E. L. Gurjeva det. (as Oed. latissimus), vid. A. Prosvirov (Moscow), HNHM coll.
Length: 3.7 – 3.9 mm (male); 4.3 – 5 mm (female), 5th abdominal segment (Fig. 2); aedeagus (Fig. 49)
Distribution: Russia (East Siberia, Far East, West Siberia), China (Heilongjiang), Kazakhstan (Cate 2007).
First record for Mongolia.

Oedostethus sincerus Gurjeva, 1975b
(Fig. 3, 4, 15, 22, 32, 33, 50-52)
subfamily Negastriinae Nakane & Kishii, 1956

Material examined. 1 ♂ - Mongolia, Zavkhan Province (Aimag), Gantsin Davaa Pass, 17 km S of Uliastai, 16.VII.1980, M. Kozlov leg., W. Dolin det. 2002, Platia coll. (Fig. 33); 2 ♂ - Mongolia, Bayankhongor Province, NW of Galuut, 2084 m a. s. l. (N46.859722° E100.113889°), 15.VII.2014, H. Bussler leg., A. Jarzabek-Müller det. 2015, A. Jarzabek-Müller coll. (Fig. 32)
Length: 3 – 3.3 mm, 5th abdominal segment (Fig. 3, 4); pronotum (Fig. 22), 2nd antennal article (Fig. 15), aedeagus (Fig. 50, 51, 52)
Distribution: Russia (Siberia) (Cate 2007)
First record for Mongolia.
 

Key to known species of the genus Oedostethus LeConte, 1853
of Mongolia (male)

1 (12). Fifth abdominal segment (=7th sternite) abruptly sinuate more or less deep at the end, the apex is extended to a more or less tapered or rounded cusp (Fig. 1-11); body flattened or slightly convex

2 (7). Second antennal segment 1.5 – 2 times longer than wide (Fig. 14, 15)

3 (4). Fifth abdominal segment very strong sinuate at the end, the apex is extended to a more or less tapered cusp; the apex of the seventh tergite is extended to a broadened and flat-topped cusp (Fig. 1); pronotum without distinctly shiny median longitudinal line; antennae extending three segments past posterior angles of pronotum; body black, antennae base and legs brown; body with grey pubescence; male genitalia with lateral denticle of paramere; length: 2.5 – 3.2 mm (female 4.1 mm); the first antennal segments as Fig. 14; aedeagus as Fig. 48.
..............................................................................Oedostethus kaszabi Gurjeva, 1968 (Fig. 30)

4 (3). Fifth abdominal segment feeble sinuate at the end, the apex is extended to a rounded cusp (Fig. 2, 3, 4)

5 (6). Size larger 3.7 – 3.9 mm (male), slightly convex; pronotum with a feeble median longitudinal line; antennae extending 1.5 – 2 segments past posterior angles of pronotum; second antennal segment 1.6 times longer than wide; male genitalia with lateral denticle of paramere; body black, antennae and legs brown; body with short grey pubescence; fifth abdominal segment as Fig. 2; aedeagus as Fig. 49; (female more convex and larger (Fig. 31); length: 4.3 – 5.0 mm).
...............................................................................Oedostethus petrenkoi Dolin, 1992 (Fig. 31)

6 (5). Size smaller 3 – 3.3 mm (male), flatter; pronotum with a broad and shiny median longitudinal line (Fig. 22); antennae extending two segments past posterior angles of pronotum; second antennal segment slightly longer than wide (Fig. 15); male genitalia without lateral denticle of paramere; body black, antennae base and legs brown; body with denser and even golden pubescence; fifth abdominal segment as Fig. 3, 4; aedeagus as Fig. 50-52.
.....................................................................Oedostethus sincerus Gurjeva, 1975b (Fig. 32, 33)

7 (2). Second antennal segment approximately as long as wide or nearly spherical (Fig. 16, 17, 18)

8 (9). Fifth abdominal segment barely perceptible sinuate at the end, the apex is wide triangular-formed (Fig. 5); body black, sometimes black-brown, more opaque; body with white pubescence; length: 4 – 5 mm; the first antennal segments as Fig. 16; aedeagus as Fig. 53.
........................................................................(Fleutiauxellus maritimus Curtis, 1840) (Fig. 34)

9 (8). Fifth abdominal segment unbowed or slightly sinuate at the end, the apex is extended to a more or less rounded or tapered cusp (Fig. 6-11)

10 (11). General body shape more robust; elytral interstriae rugose, flat and less shiny, with rough surface (Fig. 29); apex of fifth abdominal segment with rounded or more or less tapered cusp (Fig. 6-10); body black or black-brown; body with even grey pubescence; length: 3.9 – 4.8 mm (female 4.7 – 5.9 mm); the first antennal segments as Fig. 17; aedeagus as Fig. 54.
.............................................................Oedostethus latissimus Tsherepanov, 1957 (Fig. 35, 36)

11 (10). General body shape less robust and more slender, elytral interstriae less rugose and shinier, punctures in the first half of elytra feeble and sparser (Fig. 27, 28); body black with slightly gleam of bronze, sometimes dark-brown; body with grey pubescence; length: 4 – 4.5 mm; the first antennal segments as Fig. 18; aedeagus as Fig. 55.
...............................................................................Oedostethus mystax Gurjeva, 1971 (Fig. 37)

12 (1). Fifth abdominal segment regularly or slightly arcuate at the end, apex without cusp (Fig. 12, 13); body more convex

13 (16). Surface of pronotum densely granulate and more opaque, with umbilicated punctures (Fig. 23)

14 (15). Elytra with two or four yellow spots, sometimes spots are indistinct or entirely absent;
second antennal segment 2 – 2.5 times longer than wide (Fig. 19); punctures of pronotum more scattered, surface of pronotum slightly shinier (Fig. 23); elytra oval (Fig. 26); body black with gleam of ore; base of antennae, legs or only tarsi yellow; body with short yellow-grey pubescence; vary in appearance; length: 3.1 – 4 mm; aedeagus as Fig. 56.
.........................................................................Oedostethus varians Gurjeva, 1968 (Fig. 38-41)

15 (14). Elytra without yellow spots; second antennal segment 1.5 times longer than wide; punctures denser, interstices smaller than punctures, often contiguous, more opaque (except median longitudinal line); elytra with more subparallel sides (Fig. 26); body black, legs, sometimes elytra brighter; body bronze tomentose; length: 3.5 – 4.3 mm; aedeagus as Fig. 57.
................................................................Oedostethus graniger Tsherepanov, 1956 (Fig. 42-44)

16 (13). Pronotum with simple, more or less dense unumbilicated punctures, shinier (Fig. 24, 25)

17 (18). Pronotum as long as wide; body dark ore color, black or black-brown, less shiny, antennae black, legs brown-yellow, femur duller at middle, posterior angles of pronotum brighter; second antennal segment nearly two times longer than wide (Fig. 20); punctures of pronotum more dense, interstices 0.5 – 1 times larger than their diameter (Fig. 24); median longitudinal line slightly; body yellow tomentose; length: 2.6 – 3.3 mm; aedeagus as Fig. 58.
...........................................................................................Oedostethus aerarius Reitter, 1895
                                                 (syn. Oedostethus gurjevae Dolin & Bessolitzina, 1982) (Fig. 45)

18 (17). Pronotum distinctly transverse and convex; shiny black or black-brown, legs and tarsi black-brown, posterior angles of pronotum, elytral suture and epipleuron of elytron brighter; second antennal segment only 1.3 (– 1.5) times longer than wide (Fig. 21); punctures of pronotum less dense, interstices 1 – 3 times larger than their diameter (Fig. 25); smooth median longitudinal line; surface of body with very short and grey pubescence; length: 2.9 – 3.3 mm; aedeagus as Fig. 59.
......................................................................Oedostethus mediocris Gurjeva, 1972 (Fig. 46, 47)
 

Bestimmungsschlüssel der aus der Mongolei bekannten Arten der Gattung Oedostethus LeConte, 1853 (für Männchen zutreffend)

1 (12). Fünftes Abdominalsegment (=7. Sternit) mit mehr oder weniger tiefen Einbuchtungen am Ende, der Apex ist zu einem mehr oder weniger zugespitzten oder gerundetem Zipfel ausgezogen (Fig. 1-11); Körper abgeflacht oder flach gewölbt

2 (7). Zweites Fühlerglied 1.5 – 2mal länger als breit (Fig. 14, 15)

3 (4). Fünftes Abdominalsegment mit zwei tiefen Einbuchtungen am Ende, der Apex ist zu einem mehr oder weniger spitzen Zipfel ausgezogen; der Apex des siebten Tergits ist zu einem verbreiterten und abgeflachten Zipfel ausgezogen (Fig. 1); Halsschild ohne deutliche glatte Mittellinie; Fühler überragen die Halsschildhinterecken um 3 Glieder; Körper schwarz, Fühlerbasis und Beine braun; Körper mit grauer Behaarung; männliches Genital mit Haken an der Spitze der Parameren; Größe: 2.5 – 3.2 mm (Weibchen 4.1 mm); Fühlerbasis wie in Fig. 14; Aedeagus wie in Fig. 48.
..............................................................................Oedostethus kaszabi Gurjeva, 1968 (Fig. 30)

4 (3). Fünftes Abdominalsegment mit schwacher Ausbuchtung am Ende, der Apex ist zu einem abgerundeten Zipfel ausgezogen (Fig. 2, 3, 4)

5 (6). Größere Art von 3.7 – 3.9 mm Größe (Männchen), flach gewölbt; Halsschild mit schwacher Mittellinie; Fühler überragen die Halsschildhinterecken um 1.5 – 2 Glieder; zweites Fühlerglied 1. 6mal länger als breit; männliches Genital mit Haken an der Spitze der Parameren; Körper schwarz, Fühler und Beine braun; Körper mit kurzer grauer Behaarung; fünftes Abdominalsegment wie in Fig. 2; Aedeagus wie in Fig. 49; (Weibchen viel größer und gewölbter (Fig. 31); Größe: 4.3 – 5.0 mm).
...............................................................................Oedostethus petrenkoi Dolin, 1992 (Fig. 31)

6 (5). Kleinere Art von 3 - 3.3 mm Größe (Männchen), flacher; Halsschild mit ziemlich breiter glänzender und glatter Mittellinie (Fig. 22); Fühler überragen die Halsschildhinterecken um 2 Glieder; zweites Fühlerglied nur wenig länger als breit (Fig. 15); Parameren des männlichen Genitals an der Spitze ohne Haken; Körper schwarz, Fühler und Beine braun; Körper mit dichter und gleichmäßiger goldener Behaarung; fünftes Abdominalsegment wie in Fig. 3, 4; Aedeagus wie in Fig. 50-52.
.....................................................................Oedostethus sincerus Gurjeva, 1975b (Fig. 32, 33)

7 (2). Zweites Fühlerglied ungefähr so lang wie breit oder fast kugelförmig (Fig. 16, 17, 18)

8 (9). Fünftes Abdominalsegment mit kaum erkennbarer Einbuchtung, der Apex ist breit dreieckig geformt (Fig. 5); Körper schwarz, manchmal schwarz-braun, kaum glänzend; Körper mit feiner weißlicher Behaarung;
Größe 4 – 5 mm; Fühlerbasis wie in Fig. 16; Aedeagus wie in Fig. 53.
........................................................................(Fleutiauxellus maritimus Curtis, 1840) (Fig. 34)

9 (8). Fünftes Abdominalsegment gerade bis schwach eingebuchtet am Ende, der Apex zu einem mehr oder weniger gerundeten oder spitzen Zipfel geformt (Fig. 6-11)

10 (11). Körper mehr breit; Flügeldeckenzwischenräume flach und matt (Fig. 29); Apex des fünften Abdominalsegments mit einem gerundeten oder einem mehr oder weniger zugespitzten Zipfel (Fig. 6-10); Körper schwarz oder schwarz-braun; Körper mit gleichmäßiger grauer Behaarung; Größe 3.9 – 4.8 mm (Weibchen 4.7 – 5.9 mm); Fühlerbasis wie in Fig. 17; Aedeagus wie in Fig. 54.
.............................................................Oedostethus latissimus Tsherepanov, 1957 (Fig. 35, 36)

11(10). Körper schmaler; Flügeldeckenzwischenräume wenig gerunzelt und glänzender, mit wenigen kleinen Punkten besetzt, vor allem in der ersten Hälfte der Flügeldecken (Fig. 27, 28); Körper schwarz mit leichtem Bronzeschimmer, manchmal dunkelbraun; Körper grau behaart; Größe 4 – 4.5 mm; Fühlerbasis wie in Fig. 18; Aedeagus wie in Fig. 55.
..............................................................................Oedostethus mystax Gurjeva, 1971 (Fig. 37)

12 (1). Fünftes Abdominalsegment normal gerundet oder leicht eingebuchtet, das Ende nicht zu einem Zipfel geformt (Fig. 12, 13); Körper mehr konvex

13 (16). Halsschild mit dichter gekörnter und/oder genabelter Punktur (Fig. 23), etwas matt

14 (15). Flügeldecken mit zwei oder vier gelben Flecken, manchmal sind die Flecken undeutlich oder fehlen gänzlich; zweites Fühlerglied 2 – 2.5mal so lang wie breit (Fig. 19); zerstreutere Punktur des Halsschild, deshalb schwach glänzend (Fig. 23); Flügeldecken oval (Fig. 26); Körper schwarz mit Erzglanz; Fühlerbasis, Beine oder nur Tarsen gelb; Körper mit kurzer gelbgrauer Behaarung; sehr variables Aussehen; Größe 3.1 – 4 mm; Aedeagus wie in Fig. 56.
.........................................................................Oedostethus varians Gurjeva, 1968 (Fig. 38-41)

15 (14). Flügeldecken ohne gelbe Flecken; zweites Fühlerglied 1.5mal länger als breit; Punkte des Halsschilds stoßen aneinander (ausgenommen Mittellinie), deshalb Halsschild kaum glänzend; Körper paralleler (Fig. 26); Körper schwarz, Beine, manchmal auch Flügeldecken heller; Körper staubartig bronzefarben behaart; Größe 3.5 – 4.3 mm; Aedeagus wie in Fig. 57.
................................................................Oedostethus graniger Tsherepanov, 1956 (Fig. 42-44)

16 (13). Halsschild mit einfachen, mehr oder weniger dichten, ungenabelten Punkten besetzt, glänzender (Fig. 24, 25)

17 (18). Halsschild so breit wie lang; Körper dunkel erzfarben, schwarz oder schwarz-braun, matt glänzend, Fühler schwarz, Beine braun-gelb, Schenkel in der Mitte getrübt, Halsschildhinterecken heller; zweites Fühlerglied fast doppelt so lang wie breit (Fig. 20); Punktur Halsschild dicht, der Abstand zwischen den Punkten beträgt 0.5 – 1 Punktdurchmesser (Fig. 24); Halsschild mit angedeuteter Mittellinie; Körper staubartig fein gelb behaart; Größe 2.6 – 3.3 mm; Aedeagus wie in Fig. 58
...........................................................................................Oedostethus aerarius Reitter, 1895
                                                      (syn. Oedostethus gurjevae Dolin & Bessolitzina, 1982) (Fig. 45)

18 (17). Halsschild deutlich quer und konvex; schwarz glänzend oder schwarz-braun, Beine und Tarsen schwarz-braun, Halsschildhinterecken, Flügeldeckennaht und Epipleuren heller; zweites Fühlerglied nur 1.3 – (1.5mal) so lang wie breit (Fig. 21); Punktur Halsschild wenig dichter als vorhergehende Art, der Abstand zwischen den einfachen Punkten beträgt 1 – 3 Punktdurchmesser (Fig. 25); glatte Halsschildmittellinie; Oberfläche mit sehr kurzer grauer Behaarung; Größe 2.9 – 3.3 mm; Aedeagus wie in Fig. 59.
.....................................................................Oedostethus mediocris Gurjeva, 1972 (Fig. 46, 47)
 

Fig. 1: fifth abdominal segment (=7th sternite) of Oedostethus kaszabi Gurjeva (♂) (in background the seventh tergite)	Fig. 2: Dolin’s drawing of fifth abdominal segment of Oed. petrenkoi (Dolin 1992)	Fig. 3: fifth abdominal segment of Oed. sincerus Gurjeva (♂) (spec. from Bayankhongor Prov., Mongolia)
Fig. 4: fifth abdominal segment of Oedostethus sincerus Gurjeva (♂) (spec. from Zavkhan Prov., Mongolia)	Fig. 5: fifth abdominal segment of Fleutiauxellus maritimus Curtis (♂)	Fig. 6: fifth abdominal segment of Oed. latissimus Tsherep. (♂) (spec. 1)
Fig. 7: fifth abdominal segment of Oedostethus latissimus Tsherep. (♂) (spec. 2)	Fig. 8: fifth abdominal segment of Oed. latissimus Tsherep. (♂) (spec. 3)	Fig. 9: fifth abdominal segment of Oed. latissimus Tsherep. (♂) (spec. 4)
Fig. 10: fifth abdominal segment of Oedostethus latissimus Tsherep. (♂) (spec. 5)	Fig. 11: fifth abdominal segment of Oed. mystax Gurjeva (♂)	Fig. 12: fifth abdominal segment of Oed. varians Gurjeva (♂)
Fig. 13: fifth abdominal segment of Oedostethus mediocris Gurjeva (♂)	Fig. 14: the first antennal segments of Oed. kaszabi Gurj.	Fig. 15: the first antennal segments of Oed. sincerus Gurj.
Fig. 16: the first antennal segments of Fleutiauxellus maritimus Curt.	Fig. 17: the first antennal segments of Oedostethus latissimus Tsherep.	Fig. 18: the first antennal segments of Oed. mystax Gurj.
Fig. 19: the first antennal segments of Oedostethus varians Gurj.	Fig. 20: the first antennal segments of Oed. gurjevae Dol. & Bess. (=aerarius Reit.)	Fig. 21: the first antennal segments of Oed. mediocris Gurj.
Fig. 22: pronotum of Oedostethus sincerus Gurj.	Fig. 23: punctures of pronotum of Oed. varians Gurj.	Fig. 24: punctures of pronotum of Oed. gurjevae (=aerarius) Reitt.
Fig. 25: punctures of pronotum of Oedostethus mediocris Gurj.	Fig. 26: habitus and surface of Oed. varians Gurj. (♂) (left) in comparison to Oed. graniger Tsherep. (♂) (right)
Fig. 27: elytral interstriae of Oedostethus mystax Gurj.	Fig. 28: surface of elytra of Oed. mystax Gurj.	Fig. 29: surface of elytra of Oed. latissimus Tsherep.
Fig. 30: Oedostethus kaszabi Gurj. (♂) Paratype, specimen from Central aimak, Ulan-Baator, Mongolia; det. Gurjeva; coll. ZSM; length: 2.9 mm	Fig. 31: Oed. petrenkoi Dolin (♀), specimen from Uvs Province, Mongolia; det. Prosvirov; coll. HNHM; length: 6.0 mm	Fig. 32: Oed. sincerus Gurj. (♂), specimen from Bayankhongor Province, Mongolia; det. Jarzabek-Müller; coll. AJMR length: 3.0 mm
Fig. 33: Oedostethus sincerus Gurj. (♂), specimen from Zavkhan Province, Mongolia; det. Platia; coll. CPG; length: 3.06 mm	Fig. 34: Fleutiauxellus maritimus Curt. (♂), specimen from Tirol, Austria; det. Leseigneur; coll. ZSM; length: 4.3 mm	Fig. 35: Oed. latissimus Tsherep. (♂), specimen from Zavakhan Province, Mongolia; det. Jarzabek-Müller; vid. Platia; coll. AJMR; length: 4.8 mm
Fig. 36: Oedostethus latissimus Tsherep. (♂), specimen from Mongolia (Prov. unknown); det. Dolin; coll. HNHM; length: 4.5 mm	Fig. 37: Oed. mystax Gurj. (♂) Paratype, specimen from Uvs aimak, Mongolia; det. Gurjeva; coll. HNHM; length: 3.9 mm	Fig. 38: Oed. varians Gurj. (♀) Paratype, specimen from Uburchangaj aimak, Mongolia; det. Gurjeva; coll. HNHM; length: 4.0 mm
Fig. 39: Oedostethus varians Gurj. (♂), specimen from Chövsgöl aimak, Mongolia; det. Gurjeva; coll. HNHM; length: 3.2 mm	Fig. 40: Oed. varians Gurj. (♀), specimen from Uvs aimak, Mongolia; det. Gurjeva; coll. HNHM; length: 3.9 mm	Fig. 41: Oed. varians Gurj. (♂), specimen from Selenge Prov., Mongolia; det. Jarzabek-Müller; coll. AJMR; length: 3.1 mm
Fig. 42: Tsherepanov’s drawing of Oedostethus graniger (Tsherepanov 1957)	Fig. 43: Oed. graniger Tsherep. (♂), specimen from Central aimak, Ulan-Baator, Mongolia; det. Jarzabek-Müller; coll. AJMR; length: 3.3 mm	Fig. 44: Oed. graniger Tsherep. (♀), specimen from Central aimak, Ulan-Baator, Mongolia; det. Gurjeva; coll. HNHM; length: 4.1 mm
Fig. 45: Paratype of Oedostethus gurjevae Dolin & Bessolitzina, 1982 (♀), synonym of Oed. aerarius Reitter, 1895, specimen from Chitinskaya Oblast, Russia; coll. ZSM; length: 3.2 mm	Fig. 46: Oed. mediocris Gurj. (♂), specimen from Selenge Prov., Mongolia; det. Jarzabek-Müller; vid. Prosvirov; coll. AJMR; length: 2.9 mm	Fig. 47: Oed. mediocris Gurj. (♀), specimen from Selenge Prov., Mongolia; det. Jarzabek-Müller; vid. Prosvirov; coll. AJMR; length: 3.0 mm

Fig. 48: aedeagus of Oedostethus kaszabi Gurj. in ventral view; length: 0.40 mm	Fig. 49: Dolin’s drawing of aedeagus of Oed. petrenkoi (Dolin 1992)	Fig. 50: aedeagus of Oed. sincerus Gurj. in ventral view (spec. 1); length: 0.70 mm	Fig. 51: aedeagus of Oed. sincerus Gurj. in dorsal view (spec. 1); length: 0.70 mm
Fig. 52: aedeagus of Oed. sincerus Gurj. in dorsal view (spec. 2); length: 0.77 mm	Fig. 53: aedeagus of Fleutiauxellus maritimus Curt. in ventral view; length: 0.95 mm	Fig. 54: aedeagus of Oed. latissimus Tsherep. in ventral view; length: 0.98 mm	Fig. 55: aedeagus of Oed. mystax Gurj. in ventral view; length: 0.90 mm
Fig. 56: aedeagus of Oed. varians Gurj. in ventral view; length: 0.70 mm	Fig. 57: aedeagus of Oed. graniger Tsherep. in ventral view; length: 0.70 mm	Fig. 58: Dolin & Bessolitzina’s drawing of aedeagus of Oed. gurjevae (=Oed. aerarius) (Dolin & Bessolitzina, 1982)	Fig. 59: aedeagus of Oed. mediocris Gurj. in ventral view; length: 0.70 mm

Acknowledgments

     I am grateful to Alexander Prosvirov from the Moscow State University (Russia) for support in the determination of difficult species, for providing very precious information and for reading an earlier version of the manuscript.
     Also I wish to express my thanks to Giuseppe Platia (Italy) for his valuable expertise, important information and the photos of his specimen of Oedostethus sincerus.
     Thanks also to Tamás Németh and Ottó Merkl from the Hungarian Natural History Museum, Budapest (Hungary), and Michael Balke from the Bavarian State Collection of Zoology, Munich (Germany) for providing beetle specimens for this revision.

References
 

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